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Smoking had an adverse impact on both sperm concentration and total sperm counts with smokers having significantly lower sperm concentrations (median 60 versus 66 10 6 /ml, P = 0.02) and lower total sperm counts, (177 versus 226 10 6 , P 3.0.CO;2-F Harlev A, Agarwal A, Gunes SO et al (2015) Smoking and male infertility: an evidence-based review

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Importantly, the biological basis for the known age-associated accumulation of DNA methylation in Polycomb target regions remains incompletely understood